For the study of motile cells and other self-propelled particles it is often key to extract quantitative features from trajectories of these particles, both to quantify experimental trajectories and to parametrise theoretical models.

Passive particles form amorphous solids or glasses at high densities. The same is true of active particles that model living matter such as confluent tissues.

Collective tissue behaviour is inherently a multiscale phenomenon that is governed by complex biochemical and mechanical processes occurring simultaneously at the molecular, cellular, and tissue scales.

While active matter theory has successfully advanced our understanding of the collective dynamics resulting from individual sources of activity, multiple active processes usually act in concert in real biological systems.

Collective cellular activity and self-organisation phenomena arising from non-equilibrium activity are ubiquitous in tissues and cellular aggregates. However, the relationship between individual properties and biological patterns remains unexplored.